@ARTICLE{TreeBASE2Ref20006, author = {Mats Thulin and Joachim Thiede and Sigrid Liede-Schumann}, title = {Phylogeny and taxonomy of Tribulocarpus (Aizoaceae) }, year = {2012}, keywords = {Aizoaceae, molecular phylogenetics, paraphyletic species, progenitor-derivative species pairs, taxonomy, Sesuvioideae, Tetragonia, Tribulocarpus, typification}, doi = {}, url = {http://}, pmid = {}, journal = {Taxon}, volume = {61}, number = {1}, pages = {55?66}, abstract = {The phylogenetic position and circumscription of the disjunct African genus Tribulocarpus is investigated by using DNA sequences from plastid rps16 and trnL-F, and nuclear ITS. Representatives from all four subfamilies of Aizoaceae are included in the analyses, most of them from Aizooideae and Sesuvioideae. The position of Tribulocarpus as sister to the rest of Sesuvioideae is confirmed, and we agree that the genus should be referred to this subfamily. Tetragonia retusa is found to be part of the Tribulocarpus clade with strong support, and the new combination Tribulocarpus retusus is made. A lectotype is designated for Tetragonia somalensis. Tribulocarpus dimorphanthus, disjunctly distributed in south-western and north-eastern Africa, is found to be paraphyletic with regard to T. retusus, endemic in Somalia, as the latter is sister to the north-east African part of T. dimorphanthus only. Tribulocarpus dimorphanthus-T. retusus is proposed as an example of a progenitor-derivative species pair, where T. retusus has budded off from the north-eastern partial area of distribution of T. dimorphanthus. The compound and spiny fruits of T. dimorphanthus versus the simple and broadly winged fruits of T. retusus, indicate a probably rapid adaptive shift from zoochorous trample burrs to anemochorous nuts} }

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