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Citation for Study 258

About Citation title: "Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea).".
About This study was previously identified under the legacy study ID S11x16x96c21c04c59 (Status: Published).


Griswold C. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology, 539: 1-39.


  • Griswold C.


The phylogenetic relationships of spider families classically placed in the superfamily Lycosoidea, which possess a grate-shaped tapetum in some or all of the indirect eyes, are examined through exemplar taxa scored for 68 classical or newly elaborated characters. A derived calamistrum, which forms an oval to rectangular patch, is considered a synapomorphy uniting the Lycosoidea and several other genera formerly placed in the Tengellidae and Miturgidae. In the resulting analysis the Lycosoidea are shown to be monophyletic and the Tengellidae and Miturgidae polyphyletic. The Senoculidae, Oxyopidae, Stiphidiidae, and Psechridae form a monophyletic group. The monophyly of the Lycosidae, Trechaleidae (including Rhoicininae), Pisauridae plus Dolomedidae, and Zoropsidae are confirmed, whereas monophyly of the Ctenidae, Machadoniinae, and Uliodoninae are called into question.

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